The axons of the Purkinje cellsrun down through the grain cell layer and have their inhibitory termini in the cerebellar nuclei (in some cases in the vestibular nuclei). The purkinje axone emits within its proximala parts a large number of collaterals up in the molecular layer.
The Purkinje cells are excited partly by the parallel filaments of the grain cells and partly by climbing filaments from the lower olive kernel complex with great impact.
Purkinje cells are inhibited partly by themselves (proximal axon claterals) and partly by the basket and star cells.
The Purkinje cells discharge spontaneously in the frequency range 30-50/second with so-called simple action potentialser (cf. with complex spike).